CAS Ichthyology Primary Types Imagebase

SU 2003 Plagopterus argentissimus Cope [E. D.], 1874 Syntype  [Collection database record]
argentissimus, Plagopterus Cope [E. D.] 1874:130 [2] [Proceedings of the American Philosophical Society v. 14; ref. 932] "San Luis Valley, western Colorado, U.S.A." [probably southwestern Utah, U.S.A.]. Syntypes: ANSP 19851-52 (2), BMNH 1883.12.14.239 [ex. USNM 15576] (1); MCZ 18817 (2), CAS-SU 2003 (1), USNM 15576 (3). Type catalog: Böhlke 1953:36 [ref. 12291], Böhlke 1984:69 [ref. 13621], Gilbert 1998:43 [ref. 23395]. On p. 2 of separate. Miller & Hubbs 1960:34 [ref. 11934] discuss types and type locality problems. Jordan & Evermann 1896:329 [ref. 2443] listed the type as USNM 15776, but that lot contains 3 specimens. •Valid as Plagopterus argentissimus Cope 1874 -- (Lee et al. 1980:344 [ref. 22416], Page & Burr 1991:83 [ref. 18983], Swift et al. 1993:117 [ref. 22379], La Rivers 1994:475 [ref. 23389], Williams 1995:400 [ref. 22136], Sigler & Sigler 1996:107 [ref. 25943], Gilbert 1998:43 [ref. 23395], Fuller et al. 1999:138 [ref. 25838], Nelson et al. 2004:77 [ref. 27807], Scharpf 2005:36 [ref. 28940], Miller 2006:138 [ref. 28615], Minckley & Marsh 2009:158 [ref. 31114], Page & Burr 2011:181 [ref. 31215], Page et al. 2013:77 [ref. 32708], Page et al. 2023:75 [ref. 40505]). Current status: Valid as Plagopterus argentissimus Cope 1874. Cyprinidae: Plagopterinae. Distribution: Western North America: Middle and lower Colorado River system (southwestern U.S.A. and northwestern Mexico). Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 24026 Pogonichthys ciscoides Hopkirk [J. D.], 1974 Holotype  [Collection database record]
ciscoides, Pogonichthys Hopkirk [J. D.] 1974:30, Pl. 1 [University of California Publications in Zoology v. 96 (for 1973); ref. 2199] Clear Lake at Soda Bay, Lake County, California, U.S.A. Holotype: CAS 24026. Paratypes: CAS 20900 (3), 22896 (1), 22899 (1), 22907 (2), 24027-30 (6, 1, 4, 2). Additional material: CAS 18144 (6), 23906 (8), 22885 (1), 22887-88 (2, 2), 22891 (1), 22898 (1), 22900 (2); CAS-SU 33851 (6); UMMZ 197410 [ex CAS 22907] (3), 197411 [ex CAS 24029] (2). Type catalog: Gilbert 1998:61 [ref. 23395]. Appeared first (and unavailable) in Hopkirk 1968:414 [ref. 2198] without distinguishing features. •Valid as Pogonichthys ciscoides Hopkirk 1974 [but presumably extinct] -- (Lee et al. 1980:345 [ref. 22416], Page & Burr 1991:67 [ref. 18983], Gilbert 1998:61 [ref. 23395], Moyle 2002:150 [ref. 26449], Nelson et al. 2004:77 [ref. 27807], Scharpf 2005:36 [ref. 28940], Page & Burr 2011:159 [ref. 31215], Page et al. 2013:77 [ref. 32708], Page et al. 2023:75 [ref. 40505]). Current status: Valid as Pogonichthys ciscoides Hopkirk 1974. Cyprinidae: Pogonichthyinae. Distribution: Western North America: Clear Lake and tributaries, California (U.S.A.). Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 94251 Poropuntius bolovenensis Roberts [T. R.], 1998 Holotype  [Collection database record]
bolovenensis, Poropuntius Roberts [T. R.] 1998:124, Figs. 5-6 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Xe Nam Noi, 300 meters downstream from primary dam site of Xe Nam Noi-Xe-Pian, Bolovens Plateau, Sekong watershead, southern Laos. Lectotype: CAS 94251. Paralectotypes: CAS 94252-54 (110, 1, 89); plus specimens listed under the other 3 morphs. Described as a species on p. 124, as the first of 4 trinominal morphs on p. 125. Roberts states (p. 124), "In this paper intraspecific trophic morphs are formally described and named as species-level trinominals." It appears Roberts apparently attempted to make these trinominal names fall within the Code of Zoological Nomenclature; but the third name in a trinomen must not be a morph. Roberts even states on p. 125, ..only in the nominate morph or subspecies of (P. bolovenensis). These morphs are considered infrasubspecific and unavailable by Kottelat 2000:46 [ref. 25040], but they were available infrasubspecific names under the Code in force in 1998, but they apparently were not validly treated as a subspecies or species in time to be available under the 2000 edition of the Code. Lectotype established by Kottelat 2000:46 [ref. 25040] who considered all specimens of the morphs as syntypes but selected Roberts' "Holotype" for the morph bolovenensis as the lectotype. Kottelat 2000 [ref. 25040] provided new names for two of Roberts' morphs, but retained the same types. •Valid as Poropuntius bolovenensis Roberts 1998 -- (Kottelat 2000:46 [ref. 25040], Kottelat 2001:69 [ref. 25780], Kottelat 2013:143 [ref. 32989], Kang et al. 2016: 375 [ref. 40055]). Current status: Valid as Poropuntius bolovenensis Roberts 1998. Cyprinidae: Cyprininae. Distribution: Southeast Asia: Mekong River basin, southern Laos. Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 94255 Poropuntius bolovenensis acuticeps Roberts [T. R.], 1998 Holotype  [Collection database record]
acuticeps, Poropuntius bolovenensis Roberts [T. R.] 1998:125, Figs. 5-6 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Xe Nam Noi, 300 meters downstream from primary dam site of Xe Nam Noi-Xe-Pian, Bolovens Plateau, Sekong watershead, southern Laos. Holotype: CAS 94255. Paratypes: CAS 94256 (1). Described as an intraspecific trophic morph. Not available; see comments under Poropuntius bolovenensis Roberts 1998. Regarded as infrasubspecific and unavailable by Kottelat 2000:46 [ref. 25040]. Kottelat renamed this morph as the species Poropuntius consternans with the same type specimens. •Not available, infrasubspecific -- (Kottelat 2013:144 [ref. 32989] in synonymy of Poropuntius consternans). Current status: Synonym of Poropuntius consternans Kottelat 2000. Cyprinidae: Cyprininae.
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CAS 94257 Poropuntius bolovenensis glaridostoma Roberts [T. R.], 1998 Holotype  [Collection database record]
glaridostoma, Poropuntius bolovenensis morpha Roberts [T. R.] 1998:126, Figs. 5-6 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Xe Nam Noi, 300 meters downstream from primary dam site of Xe Nam Noi-Xe-Pian, Bolovens Plateau, Sekong watershead, southern Laos. Holotype: CAS 94257. Paratypes: CAS 94258-60 (10, 1, 2) Described as an intraspecific trophic morph. Not available; see comments under Poropuntius bolovenensis Roberts 1998. Regarded as infrasubspecific and unavailable by Kottelat 2000:46 [ref. 25040]. •Infrasubspecific in the synonymy of Poropuntius lobocheiloides Kottelat 2000 -- (Kottelat 2013:144 [ref. 32989]). Current status: Synonym of Poropuntius lobocheiloides Kottelat 2000. Cyprinidae: Cyprininae.
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CAS 94261 Poropuntius bolovenensis laticeps Roberts [T. R.], 1998 Holotype  [Collection database record]
laticeps, Poropuntius bolovenensis morpha Roberts [T. R.] 1998:127 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Xe Nam Noi, 300 meters downstream from primary dam site of Xe Nam Noi-Xe-Pian, Bolovens Plateau, Sekong watershead, southern Laos. Holotype (unique): CAS 94485. Described as an intraspecific trophic morph. Not available; see comments under Poropuntius bolovenensis Roberts 1998. Regarded as infrasubspecific and unavailable by Kottelat 2000:46 [ref. 25040]. Cyprinidae: Cyprininae.
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© 2001 California Academy of Sciences

CAS 94486 Poropuntius genyognathus Roberts [T. R.], 1998 Holotype  [Collection database record]
genyognathus, Poropuntius Roberts [T. R.] 1998:127, Figs. 7-8 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Chawa Kloh, upstream from Kita or Htee-tah, Tenasserim, Myanmar. Holotype: CAS 94486 (mistakenly published as CAS 94485). Paratypes: CAS 94487-93 (39, 5, 26, 9, 14, 46, 129), 94495 (3). •Valid as Poropuntius genyognathus Roberts 1998 -- (Kottelat 2013:144 [ref. 32989]). Current status: Valid as Poropuntius genyognathus Roberts 1998. Cyprinidae: Cyprininae. Distribution: Myanmar. Habitat: freshwater.
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CAS 94262 Poropuntius hathe Roberts [T. R.], 1998 Holotype  [Collection database record]
hathe, Poropuntius Roberts [T. R.] 1998:129, Fig. 9 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Rapids in Menam Moei River next to highway 1035, 129 kilometers north of Mae Sot, Thailand. Holotype: CAS 94262. Paratypes: CAS 94263-64 (11, 3), USNM 324210 (1). •Valid as Poropuntius hathe Roberts 1998 -- (Kottelat 2013:144 [ref. 32989]). Current status: Valid as Poropuntius hathe Roberts 1998. Cyprinidae: Cyprininae. Distribution: Thailand. Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 94265 Poropuntius heterolepidotus Roberts [T. R.], 1998 Holotype  [Collection database record]
heterolepidotus, Poropuntius Roberts [T. R.] 1998:129 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Nam Khong near Nam Tok Susa, Salween basin, Thailand. Holotype: CAS 94265. Paratypes: CAS 94266 (1). •Valid as Poropuntius heterolepidotus Roberts 1998 -- (Kottelat 2013:144 [ref. 32989]). Current status: Valid as Poropuntius heterolepidotus Roberts 1998. Cyprinidae: Cyprininae. Distribution: Thailand. Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 94267 Poropuntius kontumensis rasorius Roberts [T. R.], 1998 Holotype  [Collection database record]
rasorius, Poropuntius kontumensis Roberts [T. R.] 1998:130, Fig. 10 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Tributary of Da Dung and Song Dong Nai, south of Bam Ma Thuot at Cau Daktik, Sai Gon basin, Dac Lac Province, Vietnam. Holotype: CAS 94267. Paratypes: CAS 94268 (1). Roberts states, "In this paper intraspecific trophic morphs are formally described and named as species-level trinominals." They are here regarded as subspecies for purposes of conformity to the ICZN. See the account of Poropuntius bolovenensis Roberts 1998. Current status: Synonym of Poropuntius kontumensis (Chevey 1934). Cyprinidae: Cyprininae.
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© 2001 California Academy of Sciences

CAS 94269 Poropuntius melanogrammus Roberts [T. R.], 1998 Holotype  [Collection database record]
melanogrammus, Poropuntius Roberts [T. R.] 1998:130, Fig. 11 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Huay Sangkalia on road to Chedi Sam Ong, 7 kilometers north of Sangklaburi, Thailand. Holotype: CAS 94269. Paratypes: CAS 94270-71 (6, 1). •Valid as Poropuntius melanogrammus Roberts 1998 -- (Kottelat 2013:145 [ref. 32989]). Current status: Valid as Poropuntius melanogrammus Roberts 1998. Cyprinidae: Cyprininae. Distribution: Thailand. Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 94463 Poropuntius scapanognathus Roberts [T. R.], 1998 Holotype  [Collection database record]
scapanognathus, Poropuntius Roberts [T. R.] 1998:131, Fig. 12 [Natural History Bulletin of the Siam Society v. 46 (no. 1); ref. 23549] Huay Kong, just below mouth of Huay Long River, near kilometer 45 on highway 1126, northwest of Mae Hong Son, Salween basin, Thailand. Holotype: CAS 94463. Paratypes: CAS 94464-69 (2, 22, 2, 1, 56, 10); MCSNG 48357 (1). •Valid as Poropuntius scapanognathus Roberts 1998. •Synonym of Poropuntius hampaloides (Vinciguerra 1890) with question -- (Kottelat 2013:144 [ref. 32989]). Current status: Synonym of Poropuntius hampaloides (Vinciguerra 1890). Cyprinidae: Cyprininae. Distribution: Thanland (if valid). Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 501 Pseudogobio chaoi Evermann [B. W.] & Shaw [T.], 1927 Holotype  [Collection database record]
chaoi, Pseudogobio Evermann [B. W.] & Shaw [T.] 1927:106 [Proceedings of the California Academy of Sciences (Series 4) v. 16 (no. 4); ref. 1286] Nanking, Jiangsu, China. Holotype (unique): CAS-SU 501. •Synonym of Hemibarbus labeo (Pallas 1776) -- (Lu et al. 1977:444 [ref. 2850], Yue in Chen et al. 1998:238 [ref. 23878], Kottelat 2006:35 [ref. 28831], Bogutskaya et al. 2008:325 [ref. 30085], Kottelat 2013:110 [ref. 32989], Zhang et al. 2016:72 [ref. 34477], Dyldin & Orlov 2016:660 [ref. 34853]). Current status: Synonym of Hemibarbus labeo (Pallas 1776). Cyprinidae. Habitat: freshwater.
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© 2001 California Academy of Sciences

SU 23927 Ptychidio jordani Myers [G. S.], 1930 Holotype  [Collection database record]
jordani, Ptychidio Myers [G. S.] 1930:112, Fig. [Copeia 1930 (no. 4); ref. 5026] Probably Polisia or Kagi, central Taiwan. Holotype (unique): CAS-SU 23927. Type catalog: Böhlke 1953:36-37 [ref. 12291] with suggestion that type was introduced from mainland China, Ho & Shao 2011:64 [ref. 31432]. •Valid as Ptychidio jordani Myers 1930 -- (Wu et al. 1977:360 [ref. 4807], Chen & Chen 1989:378 [ref. 17650], Chu & Cui in Chu & Chen 1989:234 [ref. 13584], Zheng et al. 1989:216 [ref. 21202], Chen et al. in Pan et al. 1991:177 [ref. 23876], Zhu 1995:69 [ref. 25213], Zhang et al. in Yue 2000:229 [ref. 25272], Ho & Shao 2011:64 [ref. 31432], Zhang et al. 2016:115 [ref. 34477], Zheng et al. 2016:145 [ref. 34678]). Current status: Valid as Ptychidio jordani Myers 1930. Cyprinidae: Labeoninae. Distribution: Upper Pearl River basin, China and Vietnam (and possibly Taiwan). Habitat: freshwater.
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© 2001 California Academy of Sciences

SU 33900 Puntius binotatus banksi Herre [A. W. C. T.], 1940 Lectotype  [Collection database record]
banksi, Puntius binotatus Herre [A. W. C. T.] 1940:31 [Bulletin of the Raffles Museum No. 16; ref. 11006] 18 miles east of Kuching, Sarawak state, Borneo, East Malaysia. Lectotype: CAS-SU 33900. Paralectotypes: CAS-SU 33903 (21), ZRC 349 (20). Type catalog: Böhlke 1953:37 [ref. 12291], Alfred 1970:71 [ref. 23931]. Lectotype selected by Böhlke 1953:37 [ref. 12291]. •Synonym of Puntius binotatus (Valenciennes 1842) -- (Roberts 1989:61 [ref. 6439]). •Valid as Systomus banksi (Herre 1940) -- (Ng & Tan 1999:355 [ref. 24159], Parenti & Lim 2005:185 [ref. 28285]). •Valid as Puntius banksi Herre 1940 -- (Kottelat & Lim 1996:232 [ref. 23210], Doi 1997:11 [ref. 22832], Tan & Lim 2004:109 [ref. 27716], Rachmatika 2004:184 [ref. 28450], Grinang & Lim 2004:288 [ref. 28551], Kottelat & Widjanarti 2005:153 [ref. 28284]). •Valid as Barbodes banksi (Herre 1840) -- (Kottelat 2013:77 [ref. 32989], Sulaiman et al. 2018:26 [ref. 35802], Ng et al. 2019:521 [ref. 36606], Ren et al. 2020:[6] [ref. 37805], Sobri et al. 2021:[3] 658 [ref. 38823], Tan & Husana 2021:319 [ref. 38614], Kottelat & Lim 2021:530 [ref. 38873], Jamaluddin et al. 2022:[5] [ref. 39245]). Current status: Valid as Barbodes banksi (Herre 1940). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Borneo (Sarawak and Indonesia). Habitat: freshwater.
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© 2001 California Academy of Sciences

SU 14172 Puntius birtwistlei Herre [A. W. C. T.], 1940 Lectotype  [Collection database record]
birtwistlei, Puntius Herre [A. W. C. T.] 1940:32 [Bulletin of the Raffles Museum No. 16; ref. 11006] Chenderoh Dam, Perak, Malaysia. Lectotype: CAS-SU 14172. Paralectotypes: CAS-SU 14173 (5), ZRC 555 (1). Type catalog: Alfred 1970:71 [ref. 23931], Böhlke 1953:37 [ref. 12291]. Lectotype selected by Böhlke 1953:37 [ref. 12291]. •Valid as Poropuntius birtwistlei (Herre 1940) -- (Doi 1997:10 [ref. 22832]). •Valid as Hypsibarbus birtwistlei (Herre 1940) -- (Rainboth 1996:62 [ref. 22681], Doi 1997:6 [ref. 22832], Kottelat 2013:112 [ref. 32989], Ng et al. 2019:523 [ref. 36606]). Current status: Valid as Hypsibarbus birtwistlei (Herre 1940). Cyprinidae: Cyprininae. Distribution: Southeast Asia: Malaya (Malaysia). Habitat: freshwater.
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© 2001 California Academy of Sciences

SU 33014 Puntius kuchingensis Herre [A. W. C. T.], 1940 Holotype  [Collection database record]
kuchingensis, Puntius Herre [A. W. C. T.] 1940:11, Pl. 5 [Bulletin of the Raffles Museum No. 16; ref. 2130] 18 miles east of Kuching, Sarawak state, Borneo, East Malaysia. Holotype: CAS-SU 33014. Paratypes: CAS-SU 33015 (2). Type catalog: Böhlke 1953:37 [ref. 12291]. •Valid as Puntius kuchingensis Herre 1940 -- (Kottelat & Lim 1996:233 [ref. 23210], Doi 1997:11 [ref. 22832], Doi et al. 2001:15 [ref. 25691], Rachmatika 2004:184 [ref. 28450], Grinang & Lim 2004:289 [ref. 28551], Kottelat & Widjanarti 2005:154 [ref. 28284]). •Valid as Systomus kuchingensis (Herre 1940) -- (authors). •Valid as Barbodes kuchingensis (Herre 1940) -- (Kottelat 2013:78 [ref. 32989], Kottelat & Lim 2021:537 [ref. 38873]). Current status: Valid as Barbodes kuchingensis (Herre 1940). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Sarawak, Borneo. Habitat: freshwater.
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© 2001 California Academy of Sciences

CAS 36685 Rasbora axelrodi Brittan [M. R.], 1976 Holotype  [Collection database record]
axelrodi, Rasbora Brittan [M. R.] 1976:94, Fig. [Tropical Fish Hobbyist v. 25 (no. 4); ref. 7366] Sumatra, Indonesia (aquarium import). Holotype: CAS 36685. Paratypes: CAS 36686 (2). •Valid as Rasbora axelrodi Brittan 1976 -- (Rainboth & Kottelat 1987:419 [ref. 9123], Roberts 1989:67 [ref. 6439], Kottelat et al. 1993:44 [ref. 23448], Kottelat & Lim 1996:233 [ref. 23210], Doi 1997:12 [ref. 22832]). •Valid as Sundadanio axelrodi (Brittan 1976) -- (Fang 2003:717 [ref. 27255], Parenti & Lim 2005:185 [ref. 28285], Fang et al. 2009:17 [ref. 29983], Liao et al. 2010:155 [ref. 30521] based on placement of Sundadanio, Conway et al. 2011:259 [ref. 31697], Kottelat 2013:165 [ref. 32989], Stout et al. 2016:8 [ref. 35057] in Sundadanionidae). Current status: Valid as Sundadanio axelrodi (Brittan 1976). Cyprinidae. Distribution: Bintan Island, Sumatra. Habitat: freshwater.
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© 2001 California Academy of Sciences